Monday, November 13, 2017

The caddisfly family Uenoidae is now the caddisfly family Thremmatidae

There has been a significant change of which I've just taken notice.  In our new key for the southeast -- Larvae of the Southeastern USA: Mayfly, Stonefly, and Caddisfly Species (South Carolina, 2017) -- Uenoidae is no longer mentioned.  Uenoidae has been changed to Thremmatidae.  I'm not sure when this change occurred; I'm checking on that information.  Thremmatidae, like Uenoidae before, consists of a single genus -- Neophylax.

The keys to Thremmatidae/Neophylax ID are: "Mesonotum with anteromesal emargination; constructing stout case of rock fragments, usually with small ballast stones laterally."  (Larvae, p.292)

On the case in the picture above (Neophylax mitchelli) we can see 4 ballast stones.  The emargination on the leading edge of the mesonotum looks like this.

Another feature of note, the ventral apotome (gular sclerite) is in the form of a "T".

Our new key lists features for 16 species of Neophylax, you may recall that we've found 6 so far in our streams: N. aniqua, N. concinnus, N. consimilis, N. mitchelli, N. oligus and N. fuscus.

I've noted one thing of interest in terms of this change.  In Caddisflies: The Underwater Architects (Toronto, 2004), Wiggins had this to say: "The family Uenoidae now comprises two sub-families: Uenoinae (Farula, Neothremma, and Sericostriata of western North America  with Uenoa of Asia) and Thremmatinae (thremma of Europe, Oligophlebodes of western North America, and Neophylax widespread in North America and eastern Asia).  (Wiggins, p. 187)  It appears as though the sub-family Thremmatinae has been given full family status -- Thremmatidae.

                                                         (Neophylax consimilis)

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